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Rice was the first child of Frederick W. Smith, founder of FedEx. FedEx's first plane in 1973 was named Wendy after her.

Rice graduated from St. Mary's Episcopal School in Memphis, Tennessee. At Duke University, where she studied drama. She pursued a dramatic career in Hollywood, where she had very modest success (2 minor movie roles and 2 television episodes over a span of about 3 years). (Some of her credits are under the name "Windlind Smith".)

She married Jeffrey Scott Rice, a business executive. They had two sons, Mason Frederick Rice and Alden James Rice.

Windland Smith Rice became an accomplished photographer of nature and animals, completing commissions for organizations such as Fujifilm, the National Geographic Society, and Nature's Best Photography magazine. Her work won several awards and has been exhibited in the Smithsonian Museum of Natural History. The annual Nature’s Best Photography Windland Smith Rice Internati